Max R. Brown¹, Natacha Frachon², Edgar L. Y. Wong³, Chris Metherell⁴, Alex D. Twyford^1,2,5

¹University of Edinburgh, Institute of Evolutionary Biology, Charlotte Auerbach Road, Edinburgh, EH9 3FL, UK.

²Royal Botanic Garden Edinburgh, 20A Inverleith Row, Edinburgh, EH3 5LR, UK.

³Department of Plant Sciences, University of Oxford, South Parks Road, Oxford, OX1 3RB, UK.

⁴Botanical Society of Britain and Ireland, 4 High Firs Crescent, Harpenden, Hertfordshire, AL5 1NA.

⁵Author for correspondence (email: [email protected])

This manuscript is published in the American Journal of Botany.

The publication can now be viewed online at: https://bsapubs.onlinelibrary.wiley.com/doi/abs/10.1002/ajb2.1445

This repository contains the source code for all analyses and for figures in the main text. Included are the results of the statistical models in the "Output" folder.